Dall'individuo al superorganismo: organizzazione e vita sociale delle formiche

Convegno

Biologia e comportamento
Dalle piante all'Homo sapiens

Le colonie delle formiche, siano esse composte di milioni o da poche decine d'individui, manifestano proprietà collettive confrontabili con quelle degli organismi pluricellulari. La colonia è un "organismo" diviso in più corpi: un super organismo le cui proprietà emergenti per quanto sofisticate, derivano da meccanismi semplici e parsimoniosi che mediano le interazioni tra questi corpi. Lo studio dei principi che governano le attività collettive delle formiche, oltre a dischiuderci un mondo nascosto di mirabili eventi naturali, ha fornito agli scienziati utili conoscenze che, travalicando i confini delle discipline biologiche, si sono diffuse con grande successo anche in altri ambiti.

Prof. Donato A.Grasso
Etologo del Dipartimento di Biologia Evolutiva e Funzionale - Università degli Studi di Parma

organizzazione: Museo Civico di Rovereto - Università degli Studi di Trento CIMeC Center for Mind/Brain Sciences

eZ debug

Clear cache: Quick settings: Debug output Debug redirection Template debug Inline template debug List of used templates SQL debug output
Timing:	Apr 01 2026 01:20:53
Script start
Timing:	Apr 01 2026 01:20:53
Module start 'layout'
Timing:	Apr 01 2026 01:20:53
Module start 'content'
Timing:	Apr 01 2026 01:20:53
Module end 'content'
Debug: ezpI18n::translateText	Apr 01 2026 01:20:53
Missing translation for message in context: 'kernel/navigationpart' with comment: 'Navigation part'. The untranslated message is: 'contentclassmanager'
Debug: ezpI18n::translateText	Apr 01 2026 01:20:53
Missing translation for message in context: 'kernel/navigationpart' with comment: 'Navigation part'. The untranslated message is: 'Apps'
Debug: ezpI18n::translateText	Apr 01 2026 01:20:53
Missing translation for message in context: 'kernel/navigationpart' with comment: 'Navigation part'. The untranslated message is: 'Amministrazione Trasparente'
Debug: ezpI18n::translateText	Apr 01 2026 01:20:53
Missing translation for message in context: 'kernel/navigationpart' with comment: 'Navigation part'. The untranslated message is: 'eZTags'
Error: eZTemplate @ extension/ocbootstrap/design/ocbootstrap/templates/canonical_link.tpl:1[0]	Apr 01 2026 01:20:53
Unknown template variable 'pagedata' in namespace ''
Error: eZTemplate @ extension/ocbootstrap/design/ocbootstrap/templates/canonical_link.tpl:1[0]	Apr 01 2026 01:20:53
Unknown template variable 'pagedata' in namespace ''
Timing:	Apr 01 2026 01:20:53
Script end

Main resources:

Total runtime	0.5254 sec
Peak memory usage	28,416.0000 KB
Database Queries	114

Timing points:

Checkpoint	Start (sec)	Duration (sec)	Memory at start (KB)	Memory used (KB)
Script start	0.0000	0.0371	4,819.9063	1,695.9141
Module start 'layout'	0.0371	0.0134	6,515.8203	677.6016
Module start 'content'	0.0505	0.4387	7,193.4219	18,068.9375
Module end 'content'	0.4891	0.0361	25,262.3594	1,361.0391
Script end	0.5253		26,623.3984

Time accumulators:

Accumulator	Duration (sec)	Duration (%)	Count	Average (sec)
Ini load
Load cache	0.0272	5.1807	35	0.0008
Check MTime	0.0036	0.6827	35	0.0001
PostgreSQL Cluster
DB queries	0.0076	1.4430	8	0.0009
Postgresql Total
Database connection	0.0028	0.5286	1	0.0028
Postgresql queries	0.1730	32.9227	114	0.0015
Looping result	0.0000	0.0000	0	0.0000
MySQL Cluster
DFS operations	0.0003	0.0643	1	0.0003
TS translator
TS init	0.0087	1.6490	7	0.0012
TS cache load	0.0060	1.1496	7	0.0009
TS context load	0.0044	0.8297	7	0.0006
Template Total	0.4241	80.7	3	0.1414
Template load	0.1219	23.1990	3	0.0406
String conversion in template resource	0.0006	0.1165	26	0.0000
Template parser: create text elements	0.0134	2.5541	26	0.0005
Template parser: remove whitespace	0.0023	0.4355	26	0.0001
Template parser: construct tree	0.0745	14.1853	26	0.0029
Template load and register function	0.0104	1.9867	9	0.0012
Template processing	0.3021	57.5058	3	0.1007
states
state_id_array	0.0026	0.5035	3	0.0009
state_identifier_array	0.0036	0.6943	4	0.0009
Override
Cache load	0.0697	13.2711	39	0.0018
Sytem overhead
Fetch class attribute can translate value	0.0039	0.7396	1	0.0039
Fetch class attribute name	0.0133	2.5359	12	0.0011
XML
Image XML parsing	0.0002	0.0341	1	0.0002
class_abstraction
Instantiating content class attribute	0.0002	0.0291	12	0.0000
General
String conversion	0.0002	0.0450	32	0.0000
Note: percentages do not add up to 100% because some accumulators overlap

CSS/JS files loaded with "ezjscPacker" during request:

Cache	Type	Packlevel	SourceFiles
	CSS	0	design/base/stylesheets/core.css
	JS	0	extension/ocmaps/design/standard/javascript/leaflet.js extension/ocbootstrap/design/ocbootstrap/javascript/insertmedia.js

Templates used to render the page:

Usage	Requested template	Template	Template loaded
1	node/view/full.tpl	full/event.tpl	extension/oc_tcu/design/tcu/override/templates/full/event.tpl
1	parts/opengraph_set_persistent.tpl	<No override>	extension/ngopengraph/design/standard/templates/parts/opengraph_set_persistent.tpl
4	content/datatype/view/ezstring.tpl	<No override>	design/standard/templates/content/datatype/view/ezstring.tpl
2	content/datatype/view/ezobjectrelationlist.tpl	<No override>	extension/oc_tcu/design/tcu/templates/content/datatype/view/ezobjectrelationlist.tpl
2	content/view/embed.tpl	<No override>	extension/ocbootstrap/design/ocbootstrap/templates/content/view/embed.tpl
2	content/datatype/view/ezxmltext.tpl	<No override>	extension/ocembed/design/standard/templates/content/datatype/view/ezxmltext.tpl
2	content/datatype/view/ezxmltags/strong.tpl	<No override>	extension/ocbootstrap/design/ocbootstrap/templates/content/datatype/view/ezxmltags/strong.tpl
2	content/datatype/view/ezxmltags/line.tpl	<No override>	design/standard/templates/content/datatype/view/ezxmltags/line.tpl
1	content/datatype/view/ezxmltags/emphasize.tpl	<No override>	extension/ocbootstrap/design/ocbootstrap/templates/content/datatype/view/ezxmltags/emphasize.tpl
3	content/datatype/view/ezxmltags/paragraph.tpl	<No override>	extension/ocbootstrap/design/ocbootstrap/templates/content/datatype/view/ezxmltags/paragraph.tpl
1	parts/social_buttons.tpl	<No override>	extension/oc_tcu/design/tcu/templates/parts/social_buttons.tpl
1	parts/event/dates.tpl	<No override>	extension/oc_tcu/design/tcu/templates/parts/event/dates.tpl
1	content/datatype/view/ezemail.tpl	<No override>	extension/ocbootstrap/design/ocbootstrap/templates/content/datatype/view/ezemail.tpl
1	content/datatype/view/ezurl.tpl	<No override>	extension/ocbootstrap/design/ocbootstrap/templates/content/datatype/view/ezurl.tpl
1	atoms/meta.tpl	<No override>	extension/oc_tcu/design/tcu/templates/atoms/meta.tpl
1	print_pagelayout.tpl	<No override>	design/standard/templates/print_pagelayout.tpl
1	page_head.tpl	<No override>	extension/oc_tcu/design/tcu/templates/page_head.tpl
1	parts/opengraph_persistent.tpl	<No override>	extension/ngopengraph/design/standard/templates/parts/opengraph_persistent.tpl
1	canonical_link.tpl	<No override>	extension/ocbootstrap/design/ocbootstrap/templates/canonical_link.tpl
1	link.tpl	<No override>	extension/oc_tcu/design/tcu/templates/link.tpl
1	page_mainarea.tpl	<No override>	design/standard/templates/page_mainarea.tpl
1	page_warning.tpl	<No override>	design/standard/templates/page_warning.tpl
1	page_copyright.tpl	<No override>	design/standard/templates/page_copyright.tpl
1	setup/debug_toolbar.tpl	<No override>	design/standard/templates/setup/debug_toolbar.tpl
1	setup/clear_cache.tpl	<No override>	design/standard/templates/setup/clear_cache.tpl
1	setup/quick_settings.tpl	<No override>	design/standard/templates/setup/quick_settings.tpl
Number of times templates used: 36 Number of unique templates used: 26

Time used to render debug report: 0.0008 secs

Dall'individuo al superorganismo: organizzazione e vita sociale delle formiche

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